eChapters

Family : Musaceae

Flowering & Fruiting: Most parts of the year

Botanical description: Banana is a tropical small size tree, with large leaves which overlapping bases from the trunk. The leaves are oblong and 1.2-1.5 m in length. The stem reaches a height of about 10-30 feet. Flower develops from the centre of the crown. Only female flowers develop into a banana fruit that varies in length from 4 to12 inches. Each banana plant bears fruits only once. The plant bears flowers and fruits in most parts of the year.

Family: Nelumbonaceae

Flowering & Fruiting: July-December

Botanical description: It is a perennial aquatic plant, growing from a tuberous rootstock lying in mud at the bottom of water bodies, lakes and ponds. The plant produces large leaves which float on the surface of the water. Leaves are large, simple, glabrous, solitary, beset with scattered hard minute papillae and rounded in shape and covered with a network of microscopic hairs. Flowers are large, pink or white in colour with expanding and emerging above water. Fruit is 1-2 cm across with flat-topped seed pod divided into compartments. Seeds globose, 1 mm across and brown in colour

Family : Nymphaeaceae

Flowering: Throughout year

Fruiting: Throughout year

Botanical description: It is an aquatic perennial herb, having erect perennial root stocks that anchor it to the mud in the bottom of the water bodies. Leaves vary from being elliptic-oval to almost circular 15-26 cm across, fine hairs on upper side and always remain at floating stage. Flowers are mostly white, occasionally pink or red reaching up to 15 cm in diameter.

Family : Arecaceae

Flowering: February-March

Fruiting: April-May

Botanical Description: The plant is a moderate sized dioecious tree which grows up to 4-10 m. in height and 30 cm diameter with a large crown and rough trunk covered with persistent leaf bases. The leaves are 3 m. long, gently recurved on 1 m petioles with spines near the base. The leaf crown grows to 5 m wide and 7-10 m tall containing up to 100 leaves. The inflorescence grows upto 1 m with white flowers. The single seeded fruit ripens to orange-yellow.

Family : Phyllanthaceae

Flowering: May-June

Fruiting: June-September

Botanical Description: The plant is a small deciduous tree reaching about 25-30 ft in height. Leaves are compound, 14-25 inches long, crowded at the ends of the branches leaflets 2-3.5 inches long, alternately arranged along the rachis, ovate or obliquely ovate, acute with rounded or wedge-shaped base. Flowers are very minute, in short dense spike-like clusters arising from nodules along the branches. Fruit is pendulous, in small clusters from the branches, round or slightly flattened at the poles, with shallow or deep ribs (usually 5) 0.75 inch across.

Family : Phyllanthaceae

Flowering: March-May

Fruiting: November-February

Botanical Description: The plant is a small to medium sized deciduous tree, reaching 8 to 18 m in height. It is known for its edible fruits of the same name. The tree has crooked trunk and spreading branches. The leaves are simple, nearly stalkless and closely set along slender branchlets. The leaves are often mistaken for leaflets of pinnate leaves. The flowers are small, greenish-yellow or pinkish, having six segments, but no real petals. The fruit is nearly spherical, light greenish yellow, quite smooth and hard on appearance, with 6 vertical stripes or furrows.

Family : Mimosaceae

Flowering: January-March

Fruiting: April-June

Botanical Description: The plant is a large, nearly evergreen perennial tree that grows up to 20 m or more in height. It has a broad crown, up to 30 m across, and a short trunk, up to 1 m thick. Leaves are deciduous but foliage is persistent, as the new leaves appear while the old ones are being shed, so that the tree looks like an evergreen. Flowers yellowish-white in colour and are borne in small spherical glomerules of about 1 cm in diameter, forming short axillary panicles of 5-30 cm in length. Pods are greenish-brown to red or pinkish, rather thin with 10-15 cm long with irregular shape and flattened. There are about 10 seeds per pod, shining black in a white pulpy aril.

Family : Myrtaceae

Flowering: April-May

Fruiting: July-August

Botanical description: The plant is an evergreen tropical tree, 50 to 100 ft. tall, with oblong opposite leaves that are smooth, glossy and having a terpentine smell. It has fragrant white flowers in branched clusters at stem tips and purplish-black oval edible berries. The leaves are antibacterial, and are used for strengthning the teeth and gums. The fruit is ellipsoid or oblong, variable in size and shape, usually up to 2.5 cm long, black, shining and juicy when fully ripe. Seeds always one, with thick rounded fleshy cotyledons, closely appressed on their inner faces.

Family : Lythraceae

Flowering: August-September

Fruiting: November-December

Botanical description: The water chestnut is an annual aquatic herb with a rosette of floating leaves and submerged, anchored into the mud by very fine roots. It has two types of leaves, finely divided, feather-like submerged leaves borne along the length of the stem, and undivided floating leaves borne in a rosette at the water’s surface. The floating leaves have saw-tooth edges and are ovoid or triangular in shape, 2-3 cm long, on inflated leaf-stalks 5-9 cm long, which provide added buoyancy for the leafy portion. Flowers are white in colour, with Four petals and are borne for a long period. The fruit is a nut with barbed spines. Seeds can remain viable up to 12 years, although most germinate within the first two years.

Family : Rhamnaceae

Flowering: July-September

Fruiting: October-January

Botanical Description: Jackal Jujube is a very thorny straggling shrub with rusty-velvety young branches with paired thorns. Thorns are, one straight and the other recurved. Alternately arranged simple leaves are ovate to ovatelancelike, often oblique, with three prominent nerves and numerous transverse nervules. Tiny green flowers are borne in nearly stalkless velvety cymes in leaf axils. Fruits are spherical or obovoid drupes, black, shining, seeds woody.

1. Introduction

The remains of almond, found in several archeological sites, show the importance of this crop since centuries. Being storable after drying, the fruits can be transported everywhere, this is the reason why, these nutritious nuts have followed the human migration. The easy adaptability of this species to poor soils has favored their spread. It is an elixir for good health and a panacea for all ills. Almonds have long been an integral part of India’s cultural heritage, especially during festive seasons and weddings. In an increasingly health-conscious nation like India, the demand for almonds is ever increasing. However, several problems plague production and limit output which has led to high import.

1.1. Nutritional value

Almonds are good source of fat, in which 80% is in the form of monounsaturated fatty acids (MUFA) which decreases unwanted cholesterol of body. Almonds contain omega-6 fatty acid which is good for heart. The amount of saturated fat is very less in almonds. Almonds have high amount of phosphorus and calcium which is required for bone growth. Magnesium, manganese, vitamin B6 and Vit. E are also available in almonds. Almond oil contains glucosides, minerals, vitamins and is rich in protein. Nutritive value of almond has been presented in Table1.

1. Introduction

The cultivated apple (Malus × domestica Borkh.), is the most important fruit in temperate regions of the world. Apples were frequently used in different mythologies as symbols of immortality or reincarnation. One of the most popular stories attached to apple is that of Adam and Eve, who ate the forbidden fruit from the tree of good and evil which was in the center of the garden of Eden. The Latin word Malus has two meaning i.e. apple and evil, which has probably stemmed from this story from bible. In Greek mythology, mother earth or Gaia, presented a tree with golden apples to Zeus and his bride Hera on their wedding day. These golden apples were later indicated with many stories of love, bribery and temptation ranging from the abduction of Helen of Troy to the defeat and marriage of Atlanta. Goddess Pomona was deity of fruit trees in Rome. In many cultures, garden of paradise and pleasure is symbolized as the place with fruit-laden apple trees. Apple is popular because of its delicious taste and pleasant aroma.

1. Introduction

Apricot is among the highly nutritious fruits of Rosaceae family and appreciated for its health promoting contents. Studies have shown that apricot fruit is effective in preventing stomach ulcers and oxidative damages of living systems (Enomoto et al., 2010; Miyazawa et al., 2006). Apricot tree belongs to genus Prunus in the Rosaceae (rose) family. Apricot (Prunus armeniaca) is categorized under ‘stone fruits’, along with peaches, plums, almonds, and some cherries, due to its seed being enclosed in a hard, ‘stone’ like endocarp. This fruit is native to temperate Asia, first discovered growing wild on the mountain slopes of China, and long cultivated in Armenia later.

Throughout its history, apricot has been noted for delicious flavor, delicate velvety fruit surface, and early flowering and fruiting. The name apricot is a corruption of the Arabic and Greek al-paraecox which means the early fruit. The old spelling ‘apricock’ is retained in English until the 17^th century. The fruits are mainly consumed fresh but are also cooked and often stuffed. The high sugar content makes them suitable for drying. In china they were also preserved by salting and smoking. Other uses include fruit leathers, nectars, and liquors (Janick, 2005).

1. Introduction

Cherry is the common name of several Prunus species that include sweet cherry (P. avium), tart or sour cherry (P. cerasus), Duke cherry (hybrids between P. avium and P. cerasus). Sweet cherries are believed to have originated from the area between the Black and Caspian seas of Asia Minor. Birds probably carried the seed to Europe prior to human civilization. Cultivation was probably started by the Greeks and continued by the Romans. Its distribution and naturalization have proved extensive, for it is now found wild in areas as widely spaced as northern India, the southern plains of Europe and even on isolated islands such as Madeira. Climatic conditions are the most important limitations on cherry production. Being a deciduous tree, it needs a period of dormancy, which is initiated in temperate regions mainly by a fall in average temperatures in late autumn (Mir et al., 2017). Sweet cherry is grown almost everywhere in the temperate zone. It is a composite genus. Due to its origin, this genus needs warm and light conditions, but can also endure cold winter weather. It requires 550-600 mm precipitation annually. Sweet cherry prefers medium light (pH= 5.5–7.5), deep-layered soil, which is rich in nutrients. The root system of sweet cherry requires aeration and cannot tolerate slack water (Simon, 2004). Many areas of the world, although environmentally suited to cherry production, are currently unsuited to profitable cherry production due to high perishable nature of the fruit.

1. Introduction

Hazel nut also called as European filbert or Bhotia badam (Corylus avellana L.). Apparently the word ‘filbert’ has been derived from Middle English which in turn was a derivation of Philbert - the Saint Philibert, whose feast day in late August coincides with the ripening of the nut. Others believe that the term filbert came from the nut’s long husk being referred to as a full beard in Old English. The name “Hazel” comes from the Anglo-Saxon word for bonnet, “Haesel”; this probably derives from the greek “korys” (as in Corylus), for helmet or hood. The term “filbert” may have derived from “full beard”, descriptive of the long, leafy husk. The scientific name avellana derives from the town of Avella initially and was selected by Linnaeus from Leonhart Fuchs’s De historia stirpium commentarii insignes (1542), where the species was described as “Avellana nux sylvestris” (“wild nut of Avella”). Hazelnut is the 4^th economically important nut crop (748,000 MT/yr) of world after cashew, almond and walnut.

1. Introduction

Kiwifruit (Actinidia deliciosa Chev.) is among the most recently domesticated of all fruit plants. The kiwifruit of international commerce are large – fruited selections of Actinidia deliciosa Planch and Actinidia deliciosa (A. Chev.) C F Liang et A. R. Ferguson, belongs to family Acitinidiaceae. A. chinensis has been cultivated commercially for little more than 20 years, however, A. deliciosa for about 70 years. This is known as “China’s miracle fruit” and “Horticulture wonder of New Zealand”. Although this fruit vine originated in China, yet its full economic potential was exploited by the New Zealanders, which accounts for over 70 per cent of world trade (Franco, 1990). Kiwifruit has completely replaced the older name of ‘Chinese gooseberry’. In China, ‘Mihoutao’ (monkey peach) is used for all Actinidia species, but as A. chinensis and A. deliciosa are by far the most important economically of the various Actinidia species in China, they are often simply referred to together as Mithoutao. More explicitly, the two species are sometimes referred to together as Zhonghua (Chinese) Mihoutao. In other countries of the world, its cultivation gained momentum after 1960s and now is cultivated on a commercial scale in USA, Italy China, Japan, France, Germany and Australia.

In India, it was first introduced in 1960 at Lalbagh garden, Bangalore, but the plant did not come into bearing due to lack of chilling requirement during winter for its bud break. Later on, in 1963, it was introduced in Shimla hills, where the plant came into bearing in 1969. In order to assess the performance of kiwifruit cultivation in different agro-climatic conditions of state its plantation were done at different locations. After evaluation, this fruit was recommended for commercial cultivation in mid and low hills of the state. It can also be grown in mid hill of Jammu and Kashmir, Uttrakhand, Sikkim, Arunachal Pradesh, Meghalaya, Nagaland and Nilgiri hills of south India.

1. Introduction

Olive (Olea eurpaea L.) is an evergreen species commonly grown in the Mediterranean basin. The oil extracted from the fruit is a predominant component of the ‘Mediterranean diet’, which has documented heart- and cancer-protective benefits. These derive from the lipid composition and from biologically active molecules that accumulate during olive fruit development. The oil can reach up to 30% of the total fruit fresh weight at full ripening and is highly present in the mesocarp and at lower levels in seeds. Oil increase reaches a plateau in pulp after varaison. A marked triacylglycerol accumulation in seed and pulp occurs after endocarp lignification, when about 40 mg of oil per fruit per week can be synthesized. The fatty acid profile of the oil accumulating in the fruit is important in relation to its nutritional properties. The main fatty acid is oleic acid (C18:1), which represents about 75% of total fatty acids, followed by linoleic (C18:2), palmitic (C16:0), stearic (C18:0) and linolenic (C18:3) acids (Martinelli and Dandekar, 2014).

The olive, Olea europaea L, is the only species of the botanical family Oleaceae with edible fruit. The tree blooms with small white flowers in May, and fruit set begins almost immediately afterward in May or June. Ripening is complete toward the end of November or December, depending on the variety. Its cultivation began more than 6000 years ago in the Middle East, and spread west on both sides of the Mediterranean Sea.

1. Introduction

The peach [Prunus persica (L.) Batsch] is an important fruit crop of temperate regions, all over the world. Important centers of commercial production usually lie between latitudes 30° and 45° North and South. At higher latitudes, minimum winter temperatures and spring frosts are the limiting factors. The peach flower bud is hardy and can tolerate minimum temperature of about -23 to -26° C. Its northern range is extended where minimum temperature are ameliorated by large bodies of water, as western continental seaboards, the great Lakes in the U.S.A. and Canada and the Caspian and Black seas of Eurasia. Mid continental areas are characterized by lower winter temperatures and usually more severe spring frost hazards. Most peach cultivars require 500 to 1000 chill hours below 7.2° C to foliate and bloom. At lower altitudes, the winter requirement is not met. The peach with its smooth skin mutant is known as nectarine. Nectarines can be used in the same way as peaches, and may be considered as substitutes for peaches.

1. Introduction

Pear is among the oldest of the world’s fruit crops, figuring in both Bible and the tales of Homer. They are by far the most important of the deciduous tree fruits, are widely grown in temperate and increasingly, in tropical regions, and figure prominently in world trade. The fruit is primarily grown for the fresh fruit market, which is much more remunerative than that for processing.

Pear (Pyrus communis, Rosaceaea family), is a deciduous fruit, grown under a wide variety of climatic regimes, ranging from cold dry temperature hilly conditions to warm humid subtropical conditions. It is grown in many places of India like Jammu and Kashmir, Himachal Pradesh, Karnataka, Tamil Nadu, Kerala, Utter Pradesh. Sandy pear plants were brought to Amritsar by travellers from China who settled at Harsha Chinna village near Amritsar about 2,500 years ago. The Kashmir valley represents one of the major Pear (Pyrus communis.L.) growing areas of the world. Pear was introduced in India from China. The time at which Pear was introduced to Kashmir is not precisely known, however Yukteshwar Kumar writes in his book “A History of SinoIndian Relations” that peach and pear came to India during the early years of the Eastern Han period (25 AD to 220 AD). This pre historic fruit has been under cultivation both in Europe and Asia for about 3000 years (Hedrick et al., 1921). Between 16^th and 18^th centuries, many European writers travelling Kashmir noted fantastic royal gardens of the Mughals, mentioning the abundance of peach, pear, plum and cherry trees, with fountains spouting cold, icy water. Pear (Pyrus communis. L), originating from the German word ‘peer’ meaning fruit, is a fascinating fruit steeped in rich history. This fruit is known as ‘tang’ in Kashmiri language. Low chill requiring pears are being cultivated in plains of North India. Cultivars requiring high chilling hours (900-1000) are cultivated at higher hills of Jammu and Kashmir, Himachal Pradesh and Uttarakhand. Majority of the Pear production is actually limited to Kashmir however, low chilling cultivars grow in subtropical regions.

1. Introduction

Pecans (Carya illinoensis) are one of the most desirable trees to plant in home yards. Their size and natural beauty as a shade tree and its capacity to produce a nutritious food make them valuable. Pecan is the most widely cultivated and the most economically important member of the North American hickory group. Beyond its native range, pecan is commercially grown from the southeastern USA to the south-west (California) (Hanna, 1987, Peterson, 1990, Harlow et al., 1991). Pecan is from an Algonquian word, meaning ‘a nut requiring a stone to crack’. In Mexico, pecans and walnuts share the same Spanish name, neuz, which is a cognate of the English word nut. In the USA, pecan is considered the queen of nuts because of its value both as a wild and as a cultivated nut and acceptable quality of meats (Woodroof, 1979). Pecan is also considered as ‘queens of nuts’ because of its value both a wild and as a cultivated nut and acceptable quality of meats. As compared with other nut fruits which have been grown since time immemorial, pecan has been under cultivation only for about one hundred years but has gained enormous popularity.

1. Introduction

The pistachio nut tree, Pistacia vera L. has been extensively cultivated since ancient times in the Middle Eastern and Mediterranean countries; in India, Afghanistan. The climatic conditions favorable to pistachio culture are similar to those for almond and olives, and prevail in extensive regions of the western United States also. Pistachio is a nut which comes originally from Iran and has the history of 4000 to 5000 years. Pistachio is a drought-tolerant, slow-growing, deciduous tree that reaches to a height of 30 ft. The tree survives in arid regions where summers are hot and the winters cold. Rocky, poor terrain with little rainfall is no problem as long as there is good drainage. The fruits grow in clusters. Although called a nut, botanically, pistachios are actually drupe. The pistachio consists of a bone ivory shell that encases the edible meat portion, called the seed.

1. Introduction

Plum is one of the important stone fruit crops of temperate region throughout the world. The Japanese group of plum is native to China but was domesticated in Japan and subsequently was introduced to different parts of the world. In India, Japanese plums were introduced by European settlers and missionaries around 1870. Plum grows well in areas ranging from 900 m to 1600 m above mean sea level.

1.1. Uses

Among deciduous fruits, plum is widely distributed in the world, because of wider adaptability among different species and cultivars of plum to varied agro-climates and soil conditions. Plum is consumed as dessert, dried and processed products. Plum fruit is considered best to overcome the jaundice. Its fruit use for a preparation of juice and squash. It ranks next to peach in commercial production. It is widely cultivated in several countries in Europe, Asia and in the America.

1. Introduction

Quince is a highly nutritious underutilized fruit of sub-Himalayan region which has great economics and medicinal potential and it belongs to the group of the oldest cultural plants. Quince is known with different names, Urdu name ‘Bahee Dana’, Farsi name ‘Beh’, Greek name ‘Strythion’, Kashmiri name ‘Bamchunth’ and Hindi name ‘Bihi’ (Torkelson, 1995, Khoubnasabjafari and Jouyban, 2011). Quince is very much like pear in shape and gets a golden yellowish outer layer when mature. Quince fruit is a valuable dietary product and even then it less suitable for the direct consumption (Hricovsky et al., 2003) and for that reason they are mostly used either cooked (Alvarenga et al., 2008) or preserved (Kyzlink, 1990). However, as compared with other species, this fruit tree species is less popular throughout the world. Its predisposition to turning brown and decay is a limiting factor in the long-term storage of this fruit (Gunes, 2008). The nutritional value of the fruit is high (Bucsek et al., 1996; Kopec and Balik 2008). It is well known that quince fruit shows antioxidant effects (Silva et al., 2002) which result from the presence of a number of polyphenolic substances (Fattouch et al., 2007), e.g. flavonoids quercetin, rutin, kaemp ferol etc. (Silva et al., 2005), and also from increased levels of vitamin C (Tetera, 2006). Quince has the important nutritional and flavor composition more than many fruits. Vitamin C content of quince is nearly two times higher than in apple (Biro and Lindner, 1999).

1. Introduction

The modern cultivated strawberry (Fragaria x ananassa Duch.) is one of the most delicious, refreshing and soft fruits of the world. Worldwide it is also the most widely distributed fruit-crop due to its genotypic diversity, highly heterozygous nature and broad range of environmental adaptation. The plant is cherished in gardens and in commercial fields for its beautiful red fruit that has a tentalizing aroma (Sharma, 2002; Sharma and Sharma, 2004). Being a rich source of vitamins and minerals coupled with delicate flavour, strawberry has now become an important table-fruit of millions of people around the globe. It is amongst the few fruit crops, which gives quicker and very high returns per unit area on capital investment, as the crop is ready for harvesting within 6 months of planting. However, due to several technological advances in its cultivation like, introduction of day-neutral cultivars, and protected cultivation, strawberries remain available as fresh fruit throughout the year.

At present, strawberry is grown in wide climatic zones, extending to temperate, Mediterranean, sub-tropical and taiga zones (Sharma, 2002; Sharma and Sharma, 2004). Its cultivation is influenced by the specific regional adaptations due to critical photoperiods and temperature requirements and thus its cultural systems are highly variable. Due to the constant efforts of strawberry breeders, the world-wide interest for strawberry cultivation has boosted its production tremendously, which has resulted in widespread popularity of strawberry in the last 50 years.

1. Introduction

Persian walnut is of great importance across worldwide where nuts are harvested from wild stands, backyard gardens and rarely from commercial orchards that vary considerably in their size due to degree of management. The walnut is an old tree cultivated before the Greek age and appreciated for centuries for its fruits, wood and other parts utilized in several ways. Before the discovery of the phenomena of allelopathy, someone considered the walnut an un-healthful tree, since peasants used to wake up with migraine or fever after they had slept under a walnut tree. Some others considered the walnut a cursed tree, and in fact in Italy an old walnut tree growing in Benevento, has long been considered the place where the witches used to dance during the night of Saint John and, therefore, that tree was named “Tree of the Witches” (Avanzato, 2010).

Nuts are collected for home consumption, sale at local roadside stands and markets and for shipping to cities. Walnut trees are further utilized for their high quality timber to make a wide array of products. The leaves, bark and other plant parts are used for medicinal remedies and trees are grown and maintained for soil conservation purposes (Mamadjanov, 2006b). A diverse mix of seedlings, local selections, and advanced cultivars are grown for nut production, many with unknown origins, including some of which are the direct result of selection from the wild.

In domestic animals, skeleton (Fig. 1-1) is a framework that is formed of bones, cartilages and ligaments binding together giving the specific shape, form and support to the body. In addition, the skeletal framework surrounds the major cavities of the body giving protection to other soft visceral organs of the body. There are three major parts of skeleton in the animal body (1) the axial skeleton (2) the appendicular skeleton and (3) the splanchnic skeleton.

The axial skeleton consists of the skull, vertebral column, ribs, and sternum. The appendicular skeleton consists of the bones of fore limb and hind limb

The splanchnic skeleton includes some of the bones embedded in the substance of visceral organs in the body such as os cordis in ox and os penis in dog.

Bones

The bones forming the skeleton of the body are of different shape, size, form and the constituting tissue material. The number of different types of bones of any species may differ in various stages of the growth of the animal. So also the numbers of various bones differ from species to species.

A branch of anatomy that deals with joints or articulations in the body is termed as arthrology or syndesmology. A joint is an articulation or union formed by two or more bones or cartilages with the help of connective tissue. To form a joint, the long bones meet at their extremities e.g. joints of fore limb and hind limb. Joints of flat bones are formed by their borders or edges e.g. joint of bones of skull, the cranial and facial bones. The short bones and irregular bones form a joint by uniting at any of their parts or surfaces e.g. intervertebral joints.

Classification of joints

Joints are classified on the basis of their structure i.e. the kind of tissue material by which the bones are united, the other constituent parts that help in uniting the bones and the kind of movement performed at that joint. On this basis, the following three major types of joints exist in the skeleton of animal body.

A branch of anatomy that deals with muscles and their accessory structures is called as myology and the body system concerned with myology is called muscular system. The muscular system is composed of specialized cells called muscle fibres because of their shape in the form of fibre or spindle. The plasma of these cells contains myofibrils which are contractile elements that cause contraction of muscles in a definite manner under the influence of a stimulus which is a unique property of muscles. These cells together constitute the distinctive muscular tissue.

Classification of muscles

Based on their morphological and functional characteristics, there are three types of muscles in the body.

Smooth muscle: The muscular tissue in this type of muscles is constituted by masses of spindle shaped cells and they occur mainly in the walls of hollow visceral organs including glands, the vessels, eye ball and hair follicles. These are nonstriated muscles. Contraction in these muscles is weak but sustained and mostly consistent and rhythmic. The action of this muscle is involuntary i.e. not under the will.

Cardiac muscle: The muscle fibres of this muscle are arranged in irregular masses and they form cross striations. The ends of some fibres come in contact with the sides of others forming a kind of syncytium. This is a striated muscle and its action is involuntary. This muscle occurs only in the wall of the heart and the immediate proximal ends of the major blood vessels like aorta, venacava and the pulmonary veins.

The digestive system consists of organs concerned with prehension and digestion of food, absorption of digested food and expulsion of unabsorbed food. The digestive system extends from lips to the anus and consists of the primary and accessory organs of digestion. The primary organs include the mouth, pharynx and the alimentary canal (gastrointestinal tube) in that order of sequence. The accessory organs include the tongue, teeth, salivary glands, liver and pancreas. The alimentary canal extends from pharynx to anus and consists of the oesophagus, the stomach, the small intestine and the large intestine. The wall of the alimentary canal consists of, from within outward, the mucous membrane, the muscular layer and the serous layer forming the visceral peritoneum.

Mouth

The mouth is bounded dorsally (roof) by hard palate and behind that the soft palate, ventrally by the body of the mandible along with the mylohyoideus muscles and laterally by the cheeks. The entrance of the mouth cavity is bounded by the lips and it is called as rima oris, while its caudal opening into the pharynx is called as aditus pharyngis. The cavity of the mouth (cavum oris) is subdivided into two parts by the teeth and alveolar processes (the alveoli) of the mandible. The inner part i.e. the space within the teeth and alveoli is termed as mouth cavity proper, while the space between the cheeks and lips and the teeth and alveoli is called the vestibule of the mouth. The mucous membrane of the mouth cavity is continuous with the margin of the lips and the common integument and caudally it continues with the mucous membrane of the pharynx.

The respiratory system is a complex biological system comprised of several organs that facilitate the inhalation of oxygen and exhalation of carbon dioxide (breathing) in living organisms. The major functions of the respiratory system is providing oxygen to the body cells and removing carbon dioxide from the body. In addition to this, the respiratory system helps in controlling body temperature (especially in dogs), eliminating water (as vapour) and assisting in voice production. The organs of respiratory system extend from nose to lungs. These consist of two major parts, the upper respiratory tract and the lower respiratory tract. The upper respiratory tract is comprised of nose, nasal cavity, the part of mouth, and the part of pharynx. The lower respiratory tract is comprised of the larynx, trachea, bronchi and lungs.

Nose

The nose is the externally visible part of the respiratory system. It forms the part of the face of the animal extending from the transverse level of eyes to the rostral end of the head. Exteriorly it is formed by the bones, muscles, cartilages and the skin. Internally it encloses the nasal cavity which is lined by the mucous membrane. The dorsal part of the nose (roof of the nasal cavity) is termed as dorsum nasi. It is formed by the nasal and frontal bones of the skull. The lateral parts are formed by the incisive, maxillary, lacrimal and zygomatic bones. Covering these bones externally is a muscular layer formed by the facial muscles running over these bones and acting on the upper lip and nostrils. The muscular layer is externally covered by the skin. The skin on the nose bears fine hairs except at the tip of the nose that is formed by the nostrils. The rostral end of the nose is formed by nostrils which provide entrance to the nasal cavity. The nostrils are supported by cartilages which are the dorsal and ventral extensions of the cartilage of the nasal septum. The main aim of the nose is to support and protect the nasal cavity.

Urinary system is relatively simple and consists of a pair of kidneys, a pair of ureters, the bladder and the urethra. The kidneys are the main organs that filter the blood and produce the urine while the rests of the organs of this system are simply accessory structures that are used for transport and storage of urine. For proper functioning of the metabolic processes of the body, it is essential to constantly maintain the chemical composition and volume of the tissue fluid. This is called the homeostasis. This particular task is performed by the urinary system and in particular by the kidneys.

Kidneys

The kidneys right and left, are bean shaped red brown relatively large glands situated in the abdominal cavity. These are placed retroperitoneal against the dorsal wall of the abdomen on either side of the spine. The right kidney is about half of its length ahead of the left kidney and it fits into the renal impression on the caudate lobe of the liver. An indentation at about the middle of the medial border is the hilus of the kidney through which pass the renal vessels, nerves and ureters. Both the kidneys are covered by a thin capsule formed of fibrous tissue which surrounds the parenchyma of the kidneys. The capsule can be easily striped away from the kidney. The function of the kidneys is to maintain the volume and composition of the plasma,regulate water, ion, and pH levels, retain nutrients and excrete fluid wastes, toxins and excess electrolytes. The kidneys achieve these functions through glomerular filtration, solute reabsorption, tubular secretion, water balance and acid-base regulation.

The renal capsule provides a stiff outer shell to maintain the shape and protect the inner soft parenchyma. Deep to the capsule is the soft, dense vascular cortex. Several cone shaped pyramids form the medulla deep to the cortex. The pyramids are aligned with their bases facing outward towards the cortex and their apices point inward towards the centre of the kidney. Each apex connects to a minor calyx, a small hollow tube that collects urine. The minor calyces merge to form three major calyces, which further merge to form the hollow renal pelvis at the centre of the kidney. The urine collected into the pelvis drains into the ureter that exits through the hilus of the kidney.

Reproduction is the physiological process by which organisms produce offspring. Sexual reproduction involves production of gametes. The gametes are produced by the gonads. The rest of the process of converting the gametes into the offspring involves other reproductive organs. The male and female animals have different genital organs that contribute to reproduction. The organs of reproduction (genital organs) in male include a pair of male gonads the testes (sing.-testis), the epididymis, ductus deferens, the seminal vesicles, a prostate gland, a pair of bulbourethral glands (Cowper’s glands), the male urethra, and the penis. In female the genital organs include a pair of gonads the ovaries, the fallopian tubes, the uterus, the vagina, the vulva and the mammary glands.

Male genital system

The overall function of the male genital organs is to continuously produce and store the male gametes the sperms, to maintain the continuous supply of semen, to detect estrus in female, and to inseminate and fertilize the female gametes. While the testes are the site for production of sperms, the epididymis, vas deferens and the accessory glands like prostate, seminal vesicles, and bulbourethral glands are responsible for the production of different secretions which mix together and are known as seminal plasma or seminal fluid. The seminal fluid provides a source of energy, buffers, antioxidants and other compounds needed for sperm survivability, maturation and motility. Once spermatozoa are mixed with the seminal fluid, the combined product is called the semen.

Pituitary gland

The pituitary gland (hypophysis) lies at the base of the skull and is housed within a bony structure called the sella turcica in the body of the basisphenoid bone. It is a small pea shaped gland which is attached to the hypothalamus by the pituitary stalk. The pituitary gland has two distinct parts called the lobes. These are the anterior lobe (adenohypophysis) and the posterior lobe (neurohypophysis). The anterior lobe which is also called as anterior pituitary consists of five types of cells, each of which secretes different hormone or hormones. The types of cells, the hormones secreted by each type and the functions performed by them are narrated below.

Heart

The heart is a cone shaped thick walled and four chambered muscular organ. It lies obliquely craniocaudal in the middle mediastinum in between the left and right lungs and is covered by pericardium inside the thoracic cavity. The major part of heart (about 60 per cent) lies to the left of the middle mediastinum. The broad upper end is the base of the heart. It lies at the junction of the dorsal and middle third of the dorsoventral diameter of thorax at the level of about third to sixth rib and it faces craniodorsally. The narrow lower end is the apex. It lies just dorsal to the sternum at the level of about sixth rib and it faces caudoventrally. The surfaces, right and left are convex. They meet cranially at the right ventricular border and caudally at the left ventricular border. The right (cranial) border is strongly convex. It runs ventrally and caudally. The greater part of this border lies parallel to the sternum. The left (caudal) border is much shorter and nearly vertical. The surfaces of the heart are marked by grooves which indicate the division of the heart into four chambers the two atria dorsally and the two ventricles ventrally. The atria are separated internally by the interatrial septum. The ventricles are separated by interventricular septum. The circular coronary groove indicates the division of the heart in to atria dorsally and ventricles ventrally. It encircles the heart in its upper third part except at the level of the origin of pulmonary artery. The coronary groove is quite deep and wide. It provides passage for the coronary artery and vein. Apart from these, it is deeply filled with fat. The right (subsinuosal) interventricular groove and left (paraconal) interventricular groove indicates the line of separation between the two (right and left) ventricles by the interventricular septum. Both of these grooves run downward from the coronary groove towards the apex parallel to the respective borders and terminate just above the apex. The right and left ventricular grooves are occupied by the respective coronary vessels along with the fat.

The great vessels that leave and enter the chambers of the heart include the Aorta, the Pulmonary Artery and the Pulmonary Veins.

Aorta

The Aorta is the main systemic arterial trunk. It begins as ascending aorta at the base of the left ventricle, passes craniad and dorsad between the pulmonary artery on the left and the right atrium on the right and then turns caudad inclining slightly to the left, thus describing a curve called the aortic arch with its convexity directed upward and forward and the concavity facing downward and backward. From here it proceeds caudad on the ventrolateral surface of the thoracic vertebrae and the longus colli muscle as the descending aorta. At its origin from the base of left ventricle, the caliber of the aorta is greatest forming the bulbus aorta and here it presents three dilatations called the aortic sinuses. As the descending aorta reaches the level of the seventh thoracic vertebra it continues behind as thoracic aorta running through the middle and caudal mediastinum between the left and right lungs. The thoracic aorta first lies to the left of the median plane and then gradually inclines inwards medially and becomes median in position when it reaches to the diaphragm. It then passes through the hiatus aorticus of the diaphragm and enter the abdominal cavity as abdominal aorta. From here it passes caudad below the bodies of the lumbar vertebrae and the psoas major muscle. After reaching at the level of the sixth lumbar vertebra, the abdominal aorta terminates in the iliac quadrification by dividing into two external and two internal iliac arteries.

The lymphatic system is a part of circulatory system and a vital part of the body’s immune system. It is comprised of lymph vessels, lymph ducts, and lymph nodes that move the lymph from tissues to the blood stream and the organs such as the spleen, thymus and bone marrow that produce the lymphocytes and white blood cells. Lymph is very similar to blood plasma. During circulation through the body, the blood reaches to the capillaries at the interstitial spaces. The plasma of the blood is filtered through the capillary wall. The fresh oxygen along with nutrients, hormones and other necessary fluid matters from this plasma are released in to the interstitial fluid and about 85 per cent of the plasma along with the carbon dioxide and metabolic wastes, return into the circulation through the capillary walls. The remaining 15 per cent of the plasma along with the debris of other waste products and the lymphocytes constitutes the lymph which is returned to the circulation through an accessory route of lymphatic vessels and lymph nodes constituting the lymphatic system.

Lymph Vessels: The lymph vessels start as finger-like blind ending thin walled lymph capillaries in the interstitial connective tissue. These capillaries form a network and drain into the thicker walled collecting lymph vessels, lymph ducts and lymphatic trunks. Finally the larger lymph trunks and ducts empty their content the lymph into the cranial vena cava at the thoracic inlet. The lymph vessels contain numerous valves with one or two leaflets preventing the back flow of lymph through the lymph vessels. During its course in a centripetal direction through the lymph vessel, the lymph of each lymph vessel has to pass through at least one lymph node prior to its entry into the blood circulation. As a result there lies a chain of lymph nodes on the pathway of lymph vessels in the body.

Brain

The brain, a command centre of the body, is a complex organ. The substance of the brain is made of the grey matter and the white matter. The grey matter is often identified as cerebral cortex but other parts of the brain also contain the grey matter. Inside the brain, the grey matter is organized into large accumulations of neurons which are called the nuclei of the brain. Each nucleus is associated with a specific function. In addition to neurons, there is other type of cells the glial cells called neuroglia. The neurons are responsible for sending and receiving impulses or signals. The glial cells are nonneuronal and provide support and nutrition to neuronal cells, form myelin and facilitate signal transmission in the nervous system. The white matter is made of millions of nerve fibres that are coming from neurons and are going to establish communications with different parts of the brain and spinal cord. Since these nerve fibres are coated with myelin sheath around them these appear white and the area occupied by these nerve fibres become the white matter of the brain.

The brain has three basic components, the forebrain, midbrain and the hindbrain.

Fore brain: The forebrain (Prosencephalon) is the rostral portion of the brain. It consists of cerebrum (Telencephalon) and the thalamus, hypothalamus and pineal gland together constituting the interbrain (Diencephalon).

Mid brain: The midbrain (Mesencephalon) is the middle portion of the brain. It is in the centre lying between the interbrain and hindbrain. It consists of a cranial part of brain stem.

Hind brain: The hindbrain (Rhombencephalon) is the caudal most portion of the brain. It consists of remaining (caudal) part of the brain stem, the medulla oblongata (Myelencephalon) and the cerebellum and pons (Metencephalon).

Peripheral nerves

The peripheral nervous system (PNS) is the part of the nervous system that lies outside the brain and spinal cord. It includes cranial nerves, spinal nerves and their roots and branches, peripheral nerves and neuromuscular junctions. In the peripheral nervous system, bundles of nerve fibres or axons conduct information to and from the central nervous system. The primary role of the peripheral nervous system is to connect the central nervous system to the bones, muscles and skin of the body trunk and the limbs and the organs to allow the complex movements and behaviour.

The peripheral nervous system is divided into the somatic nervous system and the autonomic (visceral) nervous system.

The somatic nervous system is associated with skeletal muscle voluntary control of body movents. It consists of afferent and efferent nerves. Afferent nerves are concerned with relaying sensation from the body to the central nervous system. The efferent nerves are responsible for sending out commands from central nervous system to the body, stimulating muscle contraction. This system is made up of nerves that connect to the skin, sensory organs and all skeletal muscles.

The autonomic nervous system is the part of peripheral nervous system that controls the visceral functions which occur below the level of consciousness. It consists of two divisions, the sympathetic nervous system and the parasympathetic nervous system.

The sense organs are the organs of the body that respond to external stimuli by conveying impulses to the nervous system. The sense organs allow animals to sense changes in the environment around them and in their bodies so that they can respond properly. Animals can sense a wide range of stimuli that include the sense of vision, the sense of hearing, the sense of olfaction (smell), the sense of taste and the sense of touch including pain and pressure. In animals, there are two major groups of senses, the general senses and the special senses.

The bones of skull (Fig 2-1) are divided into two groups, as cranial group of bones and facial group of bones. Most of the bones of the skull are flat bones and a few are irregular bones.

Cranial bones

The cranial bones form the cranium that encloses and protects the brain and organs of special senses e.g. eye and ear. These bones include the occipital, sphenoid, ethmoid, inter-parietal, parietal, temporal and frontal. The first four of these bones are single and the rests are paired.

Occipital

The occipital is a single bone (Fig 2 -1C) and is situated on the caudo- dorsal aspect of the skull. It consists of three parts, the lateral parts are called lateral occipital, the basal part is called the basi occipital and the dorsal (squamous) part is called the squamous occipital.

Horse

The skull of horse is also pyramidal in shape like that of ox but the base of the pyramid is less wide than that of the ox. It is shorter in length and narrower in width than that of the ox. The cranium is smaller externally, however, the cavity of cranium is larger than that of the ox. The nasal cavity is divided into two chambers by nasal septum. The floor of the nasal cavity is shorter and less concave than that of the ox. The left and the right nasal chambers are completely separated and do not communicate with each other. The horn core is absent.

The occipital bone is placed at the extreme caudal part of the skull, caudal condyloid foramen is absent, the basi occipital is longer and prismatic and the squamous occipital is crossed by a nuchal crest. The sphenoid bone has two parts, the pre and post sphenoid, as that in case of ox. The wing of the post sphenoid presents three notches, carotid, oval and spinosal for internal carotid artery, ventral maxillary nerve and spheno-spinous artery respectively. The foramen orbitale and the foramen rotundum are not united. There are no major differences in the ethmoid bone of ox and horse. The frontal bones are less expansive than those of ox. These are situated between parietals caudally and nasals rostrally. The supraorbital foramen perforates the root of the supra orbital process that joins the zygomatic process of the squamous temporal. The horn core is absent. The ventral borders of the frontal bones of the opposite sides form a projection which fits into the notch formed by the two nasal bones. The temporal bone in horse has three parts, the squamous, petrous and tympanic as against two parts, the squamous and petrous of the ox. The squamous part is much more developed than that of ox. It has three parts, the body, zygomatic process and the occipital (pyramidal) process. The later one arises from the caudal part of the squamous. The stylomastoid foramen is between the hyoid and the mastoid processes and the auditory bulla is smaller. The interparietal is a single bone as that of ox.

Cervical vertebrae

The skeleton of neck (Fig 2-1) is formed by the cervical vertebrae. These are seven in number. Of these, the first and second cervical vertebrae are highly modified and deviate greatly from the general architectural plan of the vertebrae. The third, fourth and fifth cervical vertebrae are similar and confirm the general plan of the vertebrae. The sixth and seventh vertebrae have some special characteristics but do not deviate much from the general plan. The general plan of vertebrae and the vertebral column has been described in the chapter on skeletal system under Part I on systemic anatomy. Hence, only regional specific characters of the cervical vertebrae are described hereunder.

First cervical vertebra (Atlas)

The first cervical vertebra (Figure 2-2 A) is called the atlas vertebra because in man it supports the globe of the head. This is an atypical vertebra and it deviates greatly from the general plan of vertebrae. The body and the spinous process are absent. The arch forms a bony ring from which two large curved plates, the wings project laterally. The ring encloses the vertebral foramen. It forms a part of the vertebral canal through which passes the spinal cord.

Horse

The first cervical vertebra (atlas) of horse is more or less similar to that of ox with a few variations as below

The wings of atlas are curved ventrad and laterad. These are thinner than in ox but their borders are thick.

The wings present an additional foramen, the transverse foramen in the caudal part.

A faint groove on the ventral surface of the wing connects the transverse foramen to the intervertebral foramen.

Most of the bones of the skull are joined (articulated) with the adjacent bones by sutures. These are immovable joints (synarthroses). The uniting medium is a fibrous tissue and is called the sutural ligament. Most of these joints exist in the early life of the animals and as the age of the animal advances the adjacent bones fuse with each other.

The articulations of the neck are between the bones of the cervical vertebrae. The atlanto occipital articulation and the atlanto axial articulation are the special vertebral articulations while the articulations between the rest of the cervical vertebrae are common vertebral articulations.

Atlanto-occipital articulation

Type of joint: This is a condyloid type of hinge joint (Ginglymus).

Articular surfaces: The atlanto-occipital joint (Figure 2-3 B) is formed between the condyles of the occipital bone and the condyloid cavities of the atlas vertebra.

Horse

Articulations of bones of skull

Most of the joints of the bones of skull are sutures similar to those in ox. However, in the adult horse many of the sutures are effaced. The ossification of the cartilage between the basilar part of the occipital and the sphenoid occurs quite earlier. The fusion of the mandibular symphysis takes place at about second to third month after birth. The temporomandibular articulation has an additional ligament which consists of elastic tissue. It connects the post-glenoid process to the caudal border of the ramys of the mandible.

Hyoidean articulations

In the hyoidean articulations, there is only one intercornual articulation in horse instead of the proximal and distal cornual articulations appearing in the ox. The middle cornu is generally lodged in the articulation between the great and small cornu.

The muscles of the head (Fig. 2-3 A) are described in four groups viz (1) superficial muscles including the cutaneous muscle. The other muscles in this group of muscles are the muscles of lips, cheeks, nostrils, eyelids and external ear. (2) the orbital muscles (3) the mandibular muscles and (4) the hyoid muscles.

The cutaneous muscle on the head of the ox is well developed and extensive. In most of the regions of the head it is intimately adhered to the skin. The superficial fascia forms a continuous layer beneath the cutaneous muscle in this region except the natural orifices like nostrils, mouth, eyes and the external ear. The deep fascia forms a continuous layer particularly on the temporal, buccal, and the pharyngeal parts of the head. A part of the cutaneous muscle on the frontal region is called the frontalis muscle. The muscle fibres of this muscle blend with the muscles of the eyelids and the contraction of fibres of this muscle causes elevation of the superior eyelid and forms wrinkles to the skin on forehead.

Cutaneous muscle

In the cervical region of ox, the cutaneus muscle, cutaneus colli is poorly developed. It is in the form of a narrow strip on the ventral part of the neck. It arises from the ventral median connective tissue strip of the superficial fascia and extends cranially covering the cervical part of the deep fascia. The fibres of this muscle fan out dorsolaterally and form the aponeurosis with the cervical fascia and join the cutaneus facei.

Fascia

The major groups of muscles of the cervical region are covered by the thick superficial layer of fascia. The fascia is abundant and well developed in the cervical region of the ox. The fascia of the neck is in two layers, the superficial and deep. The superficial fascia is attached to the skin, whereas the deep fascia is again in two parts, the superficial and deep parts. The superficial part covers the superficial structures in the neck viz. the trapezius and brachiocephalicus muscles in the dorsolateral aspect and the jugular canal with jugular vein, and the rhomboideus muscles on the ventrolateral aspect of the neck. The superficial parts of the deep layer of cervical fascia of two sides meet at the median line both, dorsally and ventrally. The deep part of the deep fascia is attached to the vertebrae and the longus colli muscle on either side at the median line and covers the trachea, oesophagus, carotid artery etc before joining ventrally.

Horse

The cutaneous muscle in horse is thin and incompletely developed in the superficial fascia. The cutaneous muscle of the facial region (cutaneous fasciei) is incomplete and poorly developed over the mandibular space and the massetor muscle. Rostrally it passes to the angle of the mouth and blends with the orbicularis oris. The cutaneous muscle of the cervical region (cutaneous colli) is situated along the ventral region of the neck.

The frontalis muscle is absent.

The orbicularis oris forms incomplete ring around the rima oris. The levator nasolabialis is less extensive and is divided into the superior and inferior branches. Between the two branches pass the dilator naris lateralis and levator labii superioris. The levator labii superioris extends from malar, lacrimal and the maxillary bones to the upper lip in a common tendon with the muscle of the opposite side. The zygomaticus is thin, short and weak. It arises from the fascia covering the massetor. The incisivus maxillaris muscle which is absent in ox exists in horse and it lies under the mucous membrane of the upper lip. It originates on the alveolar border of the incisive bone from the level of lateral incisor to the first premolar and it inserts to the upper lip. Its action is to depress the upper lip. The incisivus mandibularis which is also absent in ox exists in horse under the mucous membrane of the lower lip. It originates at the alveolar border of the body of the mandible from the level of lateral incisor to the first premolar and is inserted on the skin of the lower lip and the chin. Its action is to raise the lower lip. The incisivus maxillaris and the incisivus mandibularis muscles acting together cause the closing of rima oris. The depressor labii inferioris is thicker and extends further back. The buccinator is almost similar to that of ox.

Horse

The fascia in the cervical region of horse has two layers, the superficial and deep. Each of these is also double layered. The superficial fascia contains the cutaneus muscle the cutaneus colli. The fascia of the two sides meets at the median line along the nuchal ligament and ventrally at the fibrous raphe.

The brachiocephalicus is undivided. It arises from the mastoid process, nuchal crest and wing of the atlas and the transverse process of the second, third and fourth cervical vertebrae. The sternocephalicus is also undivided. It arises from the cartilage of the manubrium sterni and inserts at the angle of the lower jaw. The dorsal border of this muscle forms the ventral border of the jugular furrow. The sternothyrohyoideus muscle consists of two separate but unequal parts, the sternothyroideus and the sternohyoideus. The sternothyroideus is and the sternohyoideus is larger. The first part inserts on the thyroid cartilage of the larynx and the second part inserts to the body of the hyoid bone. The omohyoideus is more extensive in horse than that in ox. It arises from the subscapular fascia near the shoulder joint and inserts at the body and the lingual process of the hyoid bone. The scalinus has two parts like those of ox, but the dorsal part in horse is smaller. It arises on the transverse process of the seventh cervical vertebra and inserts on the first rib. The ventral part is very extensive. It arises from the transverse processes of the third to sixth cervical vertebrae and inserts on the medial surface of the cranial border of the first rib. The longus capitis (rectus capitis ventralis major) arises from the transverse process of third, fourth and fifth cervical vertebrae and is inserted on the tubercle at the junction of basilar part of occipital bone with the basi sphenoid. It does not form aponeurosis either at the origin or at its insertion. The rectus capitis ventralis (rectus capitis ventralis minor) is smaller in horse than that in ox. The intertransversarii cervicis (intertransversalis colli) are smaller and present six fasciculi. The most cranial of these is separated from the common mass formed by the rest of these fascicles.